Haddad Lab Publications

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Fletcher, R. J. Jr., Betts, M. G., Damschen, E. I., Hefley, T. J., Hightower, J., Smith, T. A. H., <Fortin, M.-J., and Haddad, N. M. 2023. Addressing the problem of scale that emerges with habitat fragmentation. Global Ecology and Biogeography 32:828–841 doi.org/10.1111/geb.13658

Isbell, F., et al. 2023. Expert perspectives on global biodiversity loss and its drivers and impacts on people. Frontiers in Ecology and the Environment 21:94-103 [open access]

Kemmerling, L.R., McCarthy, A.C, Brown, C.S., Haddad, N.M. 2023. Butterfly biodiversity increases with prairie strips and conservation management in row crop agriculture. Insect Conservation and Diversity 16:828–837 doi: 10.1111/icad.12675

Resasco, J., Burt, M.A., Orrock, J.L., Haddad, N.M., Shoemaker, D., and Levey, D.J. 2023. Transient effects of corridors on polygyne fire ants over a decade. Ecological Entomology 48:263–268 doi: 10.1111/een.13214

Cayton, H.L., Haddad, N.M., Henry, E.H., Himes Boor, G.K., Kiekebusch, E.M., Morris, W.F. and Aschehoug, E.T. (2023), Restoration success varies based on time since restoration in a disturbance-dependent ephemeral wetland ecosystem. Restoration Ecology 31: e13883 doi: 10.1111/rec.13883


Kemmerling, L.R., Rutkoski, C.E., Evans, S.E., Helms IV, J.A., Cordova-Ortiz, E.S., Smith, J.D., Vázquez Custodio, J.A., Vizza, C. and Haddad, N.M., 2022. Prairie strips and lower land use intensity increase biodiversity and ecosystem services. Frontiers in Ecology and Evolution 16:828–837 doi: 10.1111/icad.12675

Burt, M. A., Resasco, J., Haddad, N. M., & Whitehead, S. R. 2022. Ants disperse seeds farther in habitat patches with corridors. Ecosphere, 13:e4324 doi: 10.1002/ecs2.4324

Schulte, L.A., Dale, B.E., Bozzetto, S., Liebman, M., Souza, G.M., Haddad, N., Richard, T.L., Basso, B., Brown, R.C., Hilbert, J.A. and Arbuckle, J.G., 2022. Meeting global challenges with regenerative agriculture producing food and energy. Nature Sustainability 5:384-388 doi: 10.1038/s41893-021-00827-y

Warneke, C. R., Caughlin, T. T., Damschen, E. I., Haddad, N. M., Levey, D. J., & Brudvig, L. A. 2022. Habitat fragmentation alters the distance of abiotic seed dispersal through edge effects and direction of dispersal. Ecology 103:e03586 [open access]

Henry, E. H., Terando, A. J., Morris, W. F., Daniels, J. C., & Haddad, N. M. 2022. Shifting precipitation regimes alter the phenology and population dynamics of low latitude ectotherms. Climate Change Ecology 100051 doi: 10.1016/j.ecochg.2022.100051

Hudson, A.R., Peters, D.P., Blair, J.M., Childers, D.L., Doran, P.T., Geil, K., Gooseff, M., Gross, K.L., Haddad, N.M., Pastore, M.A. and Rudgers, J.A. 2022. Cross-site comparisons of dryland ecosystem response to climate change in the US Long-Term Ecological Research Network. BioScience 72:889-907 doi: 10.1093/biosci/biab134

Mony, C., Uroy, L., Khalfallah, F., Haddad, N., & Vandenkoornhuyse, P. (2022). Landscape connectivity for the invisibles. Ecography 2022:e06041 doi: 10.1111/ecog.06041


Cusser, S., Haddad, N. M., & Jha, S. 2021. Unexpected functional complementarity from non-bee pollinators enhances cotton yield. Agriculture, Ecosystems & Environment 314, 107415. doi: 10.1016/j.agee.2021.107415

Griffin, S. R., & Haddad, N. M. 2021. Connectivity and edge effects increase bee colonization in an experimentally fragmented landscape. Ecography 44919-927 [open access]

Calvert, M. D., Madden, A. A., Nichols, L. M., Haddad, N. M., Lahne, J., Dunn, R. R., & McKenney, E. A. 2021. A review of sourdough starters: ecology, practices, and sensory quality with applications for baking and recommendations for future research. PeerJ 9, e11389 [open access]

Cusser, S., Pechal, J. L., & Haddad, N. M. 2021. Carrion increases pollination service across an urban gradient. Urban Ecosystems 24:243-250 doi: 10.1007/s11252-020-01032-z

Kemmerling, L. R., S. R. Griffin, and N. M. Haddad. 2021. Optimizing pollinator conservation and crop yield among perennial bioenergy crops. GCB Bioenergy 13:1030-1042 [open access]

Browning, D. M., et al. 2021. Monitoring agroecosystem productivity and phenology at a national scale: A metric assessment framework. Ecological Indicators 131:108147 [open access]

Helms IV, J. A., J. Smith, S. Clark, K. Knupp, and N. M. Haddad. 2021. Ant communities and ecosystem services in organic versus conventional agriculture in the U.S. Corn Belt. Environmental Entomology 50:1276-1285 doi: 10.1093/ee/nvab105

Cusser, S., J. A. Helms IV, C. Bahlai, and N. M. Haddad. 2021. How long do population level field experiments need to be? Utilising data from the 40-year old LTER network. Ecology Letters 24:1103-1111 [open access]

Helms IV, J. A., K. A. Roeder, S. E. Ijelu, I. Ratcliff, and N. M. Haddad. 2021. Bioenergy landscapes drive trophic shifts in generalist ants. Journal of Animal Ecology 90:738-750 [open access]


Cusser, S, Bahlai, C, Swinton, SM, Robertson, GP, and Haddad, NM. 2020. Long‐term research avoids spurious and misleading trends in sustainability attributes of no‐till. Global Change Biology 2020; 26:3715–3725. doi: 10.1111/gcb.15080

Helms IV, J. A., Ijelu, S. E., Wills, B. D., Landis, D. A., & N. M., Haddad.  2020. Ant biodiversity and ecosystem services in bioenergy landscapes. Agriculture, Ecosystems & Environment 290:106780. [open access]

Harvey, J. A., et al. 2020. International scientists formulate a roadmap for insect conservation and recovery. Nature Ecology & Evolution 4:174-176. doi: 10.1038/s41559-019-1079-8

Costanza, J. K., J. Watling, R. Sutherland, C. Belyea, B. Dilkina, H. Cayton, D. Bucklin, S. S. Romañach, and N. M. Haddad. 2020. Preserving connectivity under climate and land-use change: No one-size-fits-all approach for focal species in similar habitats. Biological Conservation 248:108678. [open access]

Henry, E. H., M. O. Burford Reiskind, A. D. Land, and N. M. Haddad. 2020. Maintaining historic disturbance regimes increases species’ resilience to catastrophic hurricanes. Global Change Biology 26:798-806. doi: 10.1111/gcb.14932


Haddad, N.M. The Last Butterflies: A Scientist’s Quest to Save a Rare and Vanishing Creature. Princeton University Press. To be published June, 2019. Available Now

Damschen, E.I., Brudvig, L.A., Burt, M.A., Fletcher, R.J., Haddad, N.M., Levey, D.J., Orrock, J.L., Resasco, J. and Tewksbury, J.J. 2019. Ongoing accumulation of plant diversity through habitat connectivity in an 18-year experiment. Science 365:1478-1480. [open access]

Chaplin-Kramer, R., R. P. Sharp, C. Weil, E. M. Bennett, U. Pascual, K. K. Arkema, K. A. Brauman, B. P. Bryant, A. D. Guerry, N. M. Haddad, M. Hamann, P. Hamel, J. A. Johnson, L. Mandle, H. M. Pereira, S. Polasky, M. Ruckelshaus, M. R. Shaw, J. M. Silver, A. L. Vogl, and G. C. Daily. 2019. Global modeling of nature’s contributions to people. Science 366:255-258. [open access]

Wepprich T, Adrion JR, Ries L, Wiedmann J, Haddad NM (2019) Butterfly abundance declines over 20 years of systematic monitoring in Ohio, USA. PLoS ONE 14(7): e0216270. [open access]

Schultz, C. B., N. M. Haddad, E. H. Henry, and E. E. Crone. 2019. Movement and Demography of At-Risk Butterflies: Building Blocks for Conservation. Annual Review of Entomology 64:167-184 [view e-print]  doi: 10.1146/annurev-ento-011118-112204

Helms, Jackson A., Selassie E. Ijelu, and Nick M. Haddad. 2019. Range expansion in an introduced social parasite-host species pair. Biological Invasions 21:2751-2759. [open access]

Caughlin, T. T., E. I. Damschen, N. M. Haddad, D. J. Levey, C. Warneke, and L. A. Brudvig. 2019. Landscape heterogeneity is key to forecasting outcomes of plant reintroduction. Ecological Applications 29:e01850. doi.org/10.1002/eap.1850

Henry, E., E. Brammer-Robbins, E. Aschehoug, and N.M. Haddad. 2019. Do substitute species help or hinder endangered species management? Biological Conservation 232:127-130. [open access]


Hawn, C. L., Herrmann, J. D., Griffin, S. R. and Haddad, N. M. 2018. Connectivity increases trophic subsidies in fragmented landscapes. Ecology Letters 21: 1620-1628. [PDF reprint] doi:10.1111/ele.12958

Fletcher, R. J., R. K. Didham, C. Banks-Leite, J. Barlow, R. M. Ewers, J. Rosindell, R. D. Holt, A. Gonzalez, R. Pardini, E. I. Damschen, F. P. L. Melo, L. Ries, J. A. Prevedello, T. Tscharntke, W. F. Laurance, T. Lovejoy, and N. M. Haddad. 2018. Is habitat fragmentation good for biodiversity? Biological Conservation 226:9-15. [PDF reprint]

Cayton, H., and N.M. Haddad. 2018. Water Availability Coincides with Population Declines for an Endangered Butterfly. Diversity 10:94. doi:10.3390/d10030094 [open online]

Haddad N.M. 2018. Resurrection and resilience of the rarest butterflies. PLoS Biology 16(2): e2003488. doi:10.1371/journal.pbio.2003488  [open online]

Herrmann, J. D., N. M. Haddad, and D. J. Levey.  2018. Mean body size predicts colony performance in the common eastern bumble bee (Bombus impatiens). Ecological Entomology 43: 458-462. 10.1111/een.12517 [open online] 


Haddad, N. M., R. D. Holt, R. J. Jr Fletcher, M. Loreau, and J. Clobert. 2017. Connecting models, data, and concepts to understand fragmentation’s ecosystem-wide effects. Ecography 40:1-8. [PDF reprint]

Haddad, N. M., Gonzalez, A., Brudvig, L. A., Burt, M. A., Levey, D. J. and Damschen, E. I. 2017. Experimental evidence does not support the Habitat Amount Hypothesis. Ecography 40:48-55. doi:10.1111/ecog.02535 [PDF reprint] 

Resasco, J., Bruna, E., Haddad, N., Banks‐Leite, C., & Margules, C. 2017. The contribution of theory and experiments to conservation in fragmented landscapes. Ecography 40:109-118 doi:10.1111/ecog.02546 [PDF reprint]

Herrmann, J.D., N.M. Haddad, and D.J. Levey. 2017. Testing the relative importance of local resources and landscape connectivity on Bombus impatiens (Hymenoptera, Apidae) colonies. Apidologie DOI: 10.1007/s13592-017-0499-1. [PDF reprint]

Haddad, N. and D. Wagner. 2017. Persistent decline in the abundance and diversity of Lepidoptera. News of the Lepidopterists’ Society 59:94-95, 101. [PDF reprint]


Sivakoff, F. S., W. F. Morris, E. T. Aschehoug, B. R. Hudgens, and N. M. Haddad. 2016. Habitat restoration alters adult butterfly morphology and potential fecundity through effects on host plant quality. Ecosphere 7(11):e01522.10.1002/ecs2.1522 [PDF reprint]

Herrmann, J.D., T.A. Carlo, L.A. Brudvig, E.I. Damschen, N.M. Haddad, D.J. Levey, J.L. Orrock, and J.J. Tewksbury.  2016.  Connectivity from a different perspective: comparing seed dispersal kernels in connected vs. unfragmented landscapes.  Ecology 1274-1282 [PDF reprint]

Levey, D.J., Caughlin, T.T., Brudvig, L.A., Haddad, N.M., Damschen, E.I., Tewksbury, J.J., Evans, D.M. 2016. Disentangling fragmentation effects on herbivory in understory plants of longleaf pine savanna.  Ecology 97:2248-2258 [PDF reprint]


Haddad, N.M. 2015. Corridors for nature, Corridors for people: How can the environmental impacts of roads be reduced? Science 350:1166-1167. [PDF reprint] [html text] [summary]

Chaplin-Kramer, R., I. Ramler, R. Sharp, N.M. Haddad, J.S. Gerber, P. West, L. Mandle, P.C. Engstrom, A. Baccini, S. Sim, C. Mueller, and H. King. 2015.  Degradation in carbon stocks near tropical forest edges.  Nature Communications 6:10158 | DOI:10.1038/ncomms10158 [PDF reprint]

Brudvig, L.A., E.I. Damschen, N.M. Haddad, D.J. Levey, and  J.J. Tewksbury. 2015. The influence of habitat fragmentation on multiple plant-animal interactions and plant reproduction.  Ecology 96:2669-2678.  [PDF reprint]

Aschehoug, E.T., F.S. Sivakoff, H.L. Cayton, W.F. Morris, and N.M. Haddad. 2015. Habitat restoration affects immature stages of a wetland butterfly through indirect effects on predation. Ecology 96:1761-1767. [PDF reprint]

Cayton, H., N.M. Haddad, K. Gross, S.E. Diamond, and L. Ries. 2015. Do growing degree days predict phenology across butterfly species? Ecology 96:1473-1479. [PDF reprint]

Cayton, H., N.M. Haddad, B. Ball, E. Henry, and E. Aschehoug. 2015. Habitat restoration as a recovery tool for a disturbance-dependent butterfly, the endangered St. Francis’ Satyr.  Pages 147-159 in Daniels, J.C. (editor) Butterfly Conservation in North America.  [PDF reprint]

Haddad, N.M., L.A. Brudvig, J. Clobert, K.F. Davies, A. Gonzalez, R.D. Holt, T.E. Lovejoy, J.O. Sexton, M.P. Austin, C.D. Collins, W.M. Cook, E.I. Damschen, R.M. Ewers, B.L. Foster, C.N. Jenkins, A.J. King, W.F. Laurance, D.J. Levey, C.R. Margules, B.A. Melbourne, A.O. Nicholls, J.L. Orrock, D.-X. Song, and J.R. Townshend. 2015. Habitat fragmentation and its lasting impact on Earth’s ecosystems. Science Advances 1, e1500052.  [PDF reprint]

Henry, E.H., N.M. Haddad, J. Wilson, P. Hughes, and B. Gardner. 2015. Point-count methods to monitor butterfly populations when traditional methods fail: a case study with Miami blue butterfly. Journal of Insect Conservation 19:519-529. [PDF reprint]

Haddad, N.M. 2015. Conservation and restoration for the endangered St. Francis’ Satyr. News of the Lepidopterists’ Society 57:33-35. [PDF reprint]

Diamond, S.E., R.R. Dunn, S.D. Frank, N.M. Haddad, and R.A. Martin. 2015. Shared and unique responses of insects to the interaction of urbanization and background climate. Current Opinion in Insect Science 11:71-77. [PDF reprint]


Breckheimer, I, N.M. Haddad, W.F. Morris, A.M. Trainor, W.R. Fields, R.T. Jobe, B.R. Hudgens, A. Moody, and J.R. Walters. 2014.  Defining and evaluating the umbrella species concept for conserving and restoring landscape connectivity. Conservation Biology 28:1584-1593. [PDF reprint]

Damschen, E.I., D.V. Baker, G. Bohrer, R. Nathan, J.L. Orrock, J.R. Turner, L.A. Brudvig, N.M. Haddad, D.J. Levey, and J.J. Tewksbury. 2014. How fragmentation and corridors affect wind dynamics and seed dispersal in open habitats. Proceedings of the National Academy of Sciences 111:3484-3489. [PDF reprint]

Resasco, J. N.M. Haddad, J.L. Orrock, D. Shoemaker, L.A. Bruvig, E.I. Damschen, J.J. Tewksbury, and D.J. Levey.  2014.  Landscape corridors can increase invasion by an exotic species and reduce diversity of native species.  Ecology 95:233-239. [PDF reprint]

Haddad, N.M., L.A. Brudvig, E.I. Damschen, D.M. Evans, B.L. Johnson, D.J. Levey, J.L. Orrock, J. Resasco, L.L. Sullivan, J.J. Tewskbury, S.A. Wagner, and A.J. Weldon.  2014. Potential negative ecological effects of corridors.  Conservation Biology 28:1178-1187. [PDF reprint]

Diamond, S.E., H. Cayton, T. Wepprich, C.N. Jenkins, R.R. Dunn, N.M. Haddad, L. Ries. 2014. Unexpected phenological responses of butterflies to the interaction of urbanization and geographic temperature.  Ecology 95:2613-2621. [PDF reprint]

Adrion, J.R., A. Kousathanas, M. Pascual, H.J. Burrack, N.M. Haddad, A.O. Bergland, H. Machado, T.B. Sackton, T.A. Schlenke, M. Watada, D. Wegmann, and N.D. Singh.  2014. Drosophila suzukii: the genetic footprint of a recent, world-wide invasion. Molecular Biology and Evolution 31:3148-3163. [PDF reprint]


Wilson, J.W., J.O. Sexton,  R.T. Jobe, and N.M. Haddad.  2013.  The relative contribution of terrain, land cover, and vegetation structure indices to species distribution models.  Biological Conservation 164:170–176. [PDF reprint]

Mata, T.M., N.M. Haddad, and M. Holyoak.  2013.  How invader traits interact with resident communities and resource availability to determine invasion success.  Oikos 122: 149–160 [PDF reprint]


Haddad, N.M.  2012.  Connecting ecology and conservation through experiment.  Nature Methods 794-795.  [PDF reprint]

Dinnage, R., M.W. Cadotte, N.M. Haddad, G.M. Crutsinger, and D. Tilman.  2012.  Diversity of plant evolutionary lineages promotes arthropod diversity.  Ecology Letters 15:1308-1317. [PDF reprint]

Hudgens, B.R., W.F. Morris, N.M. Haddad. W. Fields, J. Wilson, D.C. Kuefler, and R.T. Jobe.  2012.  How complex do models need to be to predict dispersal of threatened species through matrix habitats?  Ecological Applications 22:1701-1710.  [PDF reprint]

Milko, L.V., N.M. Haddad, and S.L. Lance.  2012.  Dispersal via stream corridors structures populations of the endangered St. Francis’ satyr butterfly (Neonympha mitchellii francisci).  Journal of Insect Conservation 16:263-273.  [PDF reprint]


Leidner, A.K. and N.M. Haddad.  2011.  Combining measures of dispersal measures to identify conservation strategies in fragmented landscapes.  Conservation Biology 25:1022-1031.  [PDF reprint]

Johnson, B.L., and N.M. Haddad. 2011. Edge effects, not connectivity, determine the incidence and development of a foliar fungal plant disease. Ecology 92:1551-1558. [PDF reprint]

Sullivan, L.L., B.L. Johnson, L.A. Brudvig, and N. Haddad. 2011. Can dispersal mode predict corridor effects on plant parasites? Ecology 92:1559-1564. [PDF reprint]

Haddad, N.M., G.M. Crutsinger, K. Gross, J. Haarstad, and D. Tilman. 2011.  Plant diversity and the stability of foodwebs.  Ecology Letters 14:42-46.  [PDF reprint]

Haddad, N.M., B. Hudgens, E.I. Damschen, D.J. Levey, J.L. Orrock, J.J. Tewksbury, and A.J. Weldon.  2011.  Assessing positive and negative ecological effects of corridors.  Pages 475-503 in J. Liu, V. Hull, A.T. Morzillo, and J.A. Wiens, Sources, sinks, and sustainability, Cambridge University Press, Cambridge, UK [PDF reprint]


Bartel, R.A., N.M. Haddad, J.P. Wright.  2010.  Ecosystem engineers maintain rare species and increase biodiversity.   Oikos 119:883-890.  [PDF reprint]

Krosby, M., J. Tewksbury, N.M. Haddad, and J. Hoesktra.  2010.  Landscape connectivity for a changing climate.  Conservation Biology 24:1686-1689.  [PDF reprint]

Kuefler, D., B. Hudgens, N.M. Haddad, W.F. Morris, and N. Thurgate. 2010.  The conflicting role of matrix habitats as conduits and barriers for dispersal.  Ecology 91:944-950. [PDF reprint]

Leidner, A.K. and N.M. Haddad.  2010.  Natural, not urban, barriers limit dispersal of a coastal endemic butterfly.  Conservation Genetics 11:2311-2320. [PDF reprint]

Leidner, A.K., Haddad, N.M., and Lovejoy, T.E.  2010.  Does tropical forest fragmentation increase long-term variability of butterfly communities?  PLoS ONE 5(3):e9534. doi:10.1371/journal.pone.0009534 [PDF reprint]


Brudvig, L.A., E.I. Damschen, J.J. Tewksbury, N.M. Haddad, and D.J. Levey.  2009.  Corridors promote biodiversity spillover into adjacent habitat.  Proceedings of the National Academy of Sciences 106:9328-9332  [PDF reprint]

Haddad, N.M., G.M. Crutsinger, K. Gross, J. Haarstad, J.M.H. Knops, and D. Tilman.  2009.  Plant species loss decreases arthropod diversity and shifts trophic structure.   Ecology Letters 12:1029-1039 [PDF reprint]  [Supplementary text]

Haddad, N.M.  2009.  Principles of reserve design.  Pages 529-537 in S. Levin, editor, Princeton Guide to Ecology.  Princeton University Press, New Jersey

Wells, C.N., R.S. Williams, G.L. Walker, and N.M. Haddad.  2009.  Effects of habitat connectivity on genetic variation of the buckeye butterfly (Junonia coenia) in a landscape experiment.  Southeastern Naturalist 8:709-722 [PDF reprint]


Damschen, E.I., L.A. Brudvig, N.M. Haddad, D.J. Levey, J.L. Orrock, and J.J. Tewksbury.  2008.  The movement ecology and dynamics of plant communities in fragmented landscapes.  Proceedings of the National Academy of Sciences 105:19078-19083.  [PDF reprint]

Haddad N.M.  2008.  Finding the corridor more traveled.  Proceedings of the National Academy of Sciences 105:19569-19570. [PDF reprint]

Haddad, N.M., B. Hudgens, C. Damiani, K. Gross, D. Kuefler, and K. Pollock.  2008.  Determining optimal monitoring for rare butterfly populations.  Conservation Biology 22:929-940.  [PDF reprint]

Haddad, N.M., M. Holyoak, T.M. Mata, K.F. Davies, B.A. Melbourne, and K. Preston.  2008.  Species’ traits predict the effects of disturbance and productivity on diversity.  Ecology Letters 11:348-356.  [PDF reprint][Supplementary text] 

Kuefler, D., N.M. Haddad, S. Hall, B. Hudgens, B. Bartel, and E. Hoffman.  2008.  Distribution, population structure, and habitat use of the endangered St. Francis’ satyr butterfly, Neonympha mitchellii francisci.  American Midland Naturalist  159:298-320.  [PDF reprint]


Gross, K., E.J. Kalendra, B.R. Hudgens, and N.M. Haddad.  2007.  Robustness and uncertainty in estimates of butterfly abundance from transect counts.  Population Ecology 49:191-200.  [PDF reprint]


Damschen, E.I., N.M. Haddad, J.L. Orrock, J.J. Tewksbury, and D.J. Levey.  2006.  Corridors increase plant species richness at large scales.  Science 313:1284-1286.  [PDF reprint]

Haddad, N.M. and J.J. Tewksbury.  2006.  Impacts of corridors on populations and communities.  Pages 390-415 in K. Crooks and M. Sanjayan, editors, Connectivity Conservation, Cambridge University Press, Cambridge, England. [PDF reprint]

Kuefler, D. and N.M. Haddad.  2006.  Local versus landscape determinants of butterfly movement behaviors. Ecography 29:549-560. [PDF reprint]


Brinkerhoff, R.J., N.M. Haddad, and J.L. Orrock.  2005.  Corridors and olfactory predator cues affect small mammal behavior.  Journal of Mammalogy 86:662-669. [PDF reprint]

Damschen, E.I., K.M. Rosenfeld, M. Wyer, D. Murphy-Medley, T.R. Wentworth, and N.M. Haddad.  2005. Visibility matters: increasing knowledge of women’s contributions to ecology.  Frontiers in Ecology and the Environment 3:212-219.  [PDF reprint]

Haddad, N.M. 2005. Butterflies of the Savannah River Site. Pp. 175-184. In J. Kilgo and J. Blake, eds, Ecology and Management of a Forested Landscape, Island Press, Washington, DC.

Haddad, N.M. and J.J. Tewksbury.  2005.  Low quality habitat corridors as movement conduits for two butterfly species.  Ecological Applications 15:250-257.  [PDF reprint]

Levey, D.J., B.M. Bolker, J.J. Tewksbury, S. Sargent, and N.M. Haddad.  2005.  Effects of landscape corridors on seed dispersal by birds.  Science 309:146-148.   [PDF reprint]  [review article in Science]

Levey, D.J., B.M. Bolker, J.J. Tewksbury, S. Sargent, and N.M. Haddad.  2005.  Landscape corridors: Possible dangers? (response).  Science 310:779-783.  [PDF reprint]

Weldon, A.J. and N.M. Haddad.  2005.  The effects of patch shape on Indigo Buntings: evidence for an ecological trap.  Ecology 86:1422-1431. [PDF reprint]


Bradley, K.L., E.I. Damschen, L.M. Young, D. Kuefler, S. Went, G. Wray, N.M. Haddad, J.M.H. Knops, and S.M. Louda.  2003.  Spatial heterogeneity, not visitation bias, dominates variation in herbivory. Ecology 84:2214-2221. [PDF reprint]

Haddad, N.M., D.R. Bowne, A. Cunningham, B. Danielson, D. Levey, S. Sargent, and T. Spira.  2003.  Corridor use by diverse taxa.  Ecology 84:609-615. [PDF reprint]

Hudgens, B.R. and N.M. Haddad.  2003.  Predicting which species will benefit from corridors in fragmented landscapes from population growth models.  The American Naturalist 161:808-820.  [PDF reprint]


Haddad, N.M., D. Tilman, and J.M.H. Knops.  2002.  Long-term oscillations in grassland productivity induced by drought.  Ecology Letters 5:110-120. [PDF reprint]

Sisk, T.D., and N.M. Haddad.  2002.  Incorporating the effects of habitat edges into landscape models:  Effective area models for management.  Pp. 208-240 in J. Liu and W.W. Taylor, Integrating landscape ecology into natural resource management, Cambridge University Press, Cambridge, UK.

Tewksbury, J.J., D.J. Levey, N.M. Haddad, S. Sargent, J.L. Orrock, A. Weldon, B.J. Danielson, J. Brinkerhoff, E.I. Damschen, and P. Townsend.  2002.  Corridors affect plants, animals, and their interactions in fragmented landscapes. Proceedings of the National Academy of Sciences 99:12923-12926.  Outstanding Paper Award in Landscape Ecology [PDF reprint]


Haddad, N.M., D. Tilman, J. Haarstad, M. Ritchie, and J. Knops.  2001.  Contrasting effects of plant richness and composition on insect communities: a field experiment.  The American Naturalist 158:17-35. [PDF reprint]


Haddad, N.M. 2000. Corridor length and patch colonization by a butterfly, Junonia coeniaConservation Biology 14:738-745. [PDF reprint]

Haddad, N.M., D.K. Rosenberg, and B.R. Noon. 2000. On experimentation and the study of corridors.  Conservation Biology 14:1543-1545. [PDF reprint]

Haddad, N.M., J. Haarstad, and D. Tilman. 2000. The effects of long-term nutrient loading on grassland insect communities. Oecologia 124:73-84. [PDF reprint]

Haddad, N.M. and W.M. Hicks. 2000. Host pubescence and the behavior and performance of a butterfly, Papilio troilus (Lepidoptera). Environmental Entomology 29:299-303.  [PDF reprint]


Haddad, N.M. 1999. Corridor use predicted from behaviors at habitat boundaries. The American Naturalist 153:215-227. [PDF reprint]

Haddad, N.M. 1999. Corridor and distance effects on interpatch movements: a landscape experiment with butterflies. Ecological Applications 9:612-622.  [PDF reprint]

Haddad, N.M. and K. Baum. 1999. An experimental test of corridor effects on butterfly densities. Ecological Applications 9:623-633.  [PDF reprint]

Knops, J.M.H., D. Tilman, N.M. Haddad, S. Naeem, C.E. Mitchell, J. Haarstad, M.E. Ritchie, K.M. Howe, P.B. Reich, E. Siemann, and J. Groth. 1999.  Cascading effects of plant diversity on invasions, diseases, and insects. Ecology Letters 2:286-293. [PDF reprint]


Sisk, T.D., N.M. Haddad, and P.R. Ehrlich. 1997. Bird assemblages in patchy woodlands: modeling the effects of edge and matrix habitats.  Ecological Applications 7:1170-1180.  [PDF reprint]

Austin, G.T., N.M. Haddad, C.A. Mendez, A.E. Launer, and P.R. Ehrlich. 1996. Annotated checklist of the butterflies of the Tikal National Park area of Guatemala. Tropical Lepidoptera 7:21-37. [PDF reprint]

Mendez, C.A., T.D. Sisk, and N.M. Haddad.  1995.  Beyond birds: multitaxonomic monitoring programs provide a broad measure of tropical diversity. Pages 451-456 in J.A. Bissonette and P.R. Krausman, eds. Integrating people and wildlife for a sustainable future. Proceedings of the First International Wildlife Management Congress. The Wildlife Society, Bethesda, MD.

Pulliam, H.R. and N.M. Haddad. 1994. Human population growth and the carrying capacity concept. Bulletin of the Ecological Society of America. 75:141-157. [PDF reprint]

Daily, G.C., P.R. Ehrlich, and N.M. Haddad. 1993. Double keystone bird in a keystone species complex. Proceedings of the National Academy of Sciences 90:592-594.  [PDF reprint]